PA Natural Heritage Program Species Lists PFBC, PGC & DCNR .
PA Natural Heritage Program Species ListsPFBC, PGC & DCNREndangered, Threatened, Special Concern Species & Rare and Significant Ecological FeaturesList as of Jan 28, 2014This document provides lists of Threatened and Endangered species, Special Concern Species, and Rare and Significant Ecological features in Pennsylvania that are found in the PA Natural Diversity Inventory(PNDI) database and tool. The species and resources have been categorized according to the PA Department of Environmental Protection’s definitions in their PNDI Policy. Species and resources are listed byeach jurisdictional agency or commission. Each agency or commission provides recommendations for their species or resources to landowners, developers, and planners to conserve or enhance the speciesthrough the PNDI review process.For guidance on how PNDI recommendations are implemented when a DEP permit is required, please see the PA Department of Environmental Protection’s (DEP) PNDI et/Document-95198/021-0200-001.pdf).Also, please visit the PNHP website for more information on all species and ecological features surveyed and tracked by the PA Natural Heritage Program ons for each agency or commission Rank and Status can be found at the end of this document.PA Fish & Boat CommissionTotal Reptiles, Amphibians, Fish and Mussels in PAReptilesAmphibiansFish3736160(217)*includes species that visit PA, but not year round residentMusselsNon-mussel Aquatic InvertebratesTotal (excluding non-mussel aquatic invertebrates)65 10,000298 (355*)T&E speciesThreatenedEndangeredTOTAL144862(20% of all reptiles, amphibians, fish and mussels)Special Concern SpeciesCandidateOther SpeciesTOTAL104454(17% of all reptiles, amphibians, fish and mussels)Page 1 of 35
PA Game CommissionTotal Birds and Mammals in PABirds186(414)*includes species that have been found in PA, ofwhich 286 regularly (annually) occurMammalsTotal66252 (480)T&E SpeciesThreatenedEndangeredTOTAL71926(10% of resident birds and mammals)Special Concern Species and Rare and Significant Ecological ResourcesOther SpeciesWinter Bat ColonyTOTAL14115(5% of resident birds and mammals)Page 2 of 35
PA Department of Conservation and Natural ResourcesTotal Native Plants in PANative PlantsRare and Significant Ecological Features3,348415 tracked features by PNHPT&E SpeciesThreatened plantsEndangered plants78228306TOTAL(9% of all native plants)Special Concern SpeciesRare plantsProposed T& E plantsProposed Rare plants3918420243TotalRare and Significant Ecological FeaturesTentatively Undetermined plantsonly, not TU with a Proposed status)Natural CommunitiesInvertebrates (moths, butterflies)Geologic featuresTOTAL(7% of all native plants)(TU6556(PA has 147 classified natural communities)51(PA has 12,000 native terrestrial invertebrates)18195Page 3 of 35
PA Natural Heritage Program Species ListsReptiles, Amphibians, Fish and Aquatic Species--PA Fish and Boat CommissionThreatened and Endangered Species & Species of ConcernList as of Jan 28, 2014Reptiles, Amphibians, Fish and Aquatic Threatened and Endangered 27282930313233343536Scientific NameAcipenser brevirostrumAcipenser fulvescensAcipenser oxyrinchusAcris crepitansAlasmidonta heterodonAlosa mediocrisAmbystoma lateraleAmeiurus melasCatostomus catostomusChaenobryttus gulosusClonophis kirtlandiiCoregonus artediEnneacanthus obesusEpioblasma torulosa rangianaEpioblasma triquetraErimystax x-punctatusEtheostoma exileEtheostoma pellucidaGasterosteus aculeatusGlyptemys muhlenbergiiIchthyomyzon fossorIctiobus cyprinellusKinosternon subrubrum subrubrumLepisosteus oculatusLepomis megalotisLithobates sphenocephalus utriculariusLota lotaLythrurus umbratilisMargaritifera margaritiferaNotropis bifrenatusNotropis blenniusNotropis buchananiNotropis chalybaeusNotropis heterodonNoturus eleutherusNoturus gyrinusCommon NameShortnose SturgeonLake SturgeonAtlantic SturgeonNorthern Cricket FrogDwarf WedgemusselHickory ShadBlue-spotted SalamanderBlack BullheadLongnose SuckerWarmouthKirtland's SnakeCiscoBanded SunfishNorthern RiffleshellSnuffboxGravel ChubIowa DarterEastern Sand DarterThreespine SticklebackBog TurtleNorthern Brook LampreyBigmouth BuffaloEastern Mud TurtleSpotted GarLongear SunfishSouthern Leopard FrogBurbotRedfin ShinerEastern PearlshellBridle ShinerRiver ShinerGhost ShinerIroncolor ShinerBlackchin ShinerMountain MadtomTadpole MadtomPage 4 of 35Global G4G5G5T5G5G5G5TNRG5G5G4G3G5G5G4G5G4G5State S1S1S1S3S1S1S1S1S1S1S1S1S1Status PEPEPEPEPEPEPEPEPEPEPEPEPE
62Noturus stigmosusObovaria subrotundaOpheodrys aestivusPhoxinus eosPleurobema clavaPseudacris kalmiPseudotriton montanus montanusQuadrula cylindricaSimpsonaias ambiguaSistrurus catenatus catenatusTritogonia verrucosaVillosa fabalisAneides aeneusEtheostoma camurumEtheostoma maculatumEtheostoma tippecanoeIchthyomyzon greeleyiMinytrema melanopsNotropis dorsalisNoturus miurusPercina bimaculataPercina evidesPhoxinus erythrogasterPlethobasus cyphyusPseudemys rubriventrisScaphiopus holbrookiiNorthern MadtomRound HickorynutRough Green SnakeNorthern Redbelly DaceClubshellNew Jersey Chorus FrogEastern Mud SalamanderRabbitsfootSalamander MusselEastern MassasaugaPistolgrip MusselRayed Bean MusselGreen SalamanderBluebreast DarterSpotted DarterTippecanoe DarterMountain Brook LampreySpotted SuckerBigmouth ShinerBrindled MadtomChesapeake LogperchGilt DarterSouthern Redbelly DaceSheepnose MusselEastern Redbelly TurtleEastern SpadefootPage 5 of PEPEPEPEPEPEPEPTPTPTPTPTPTPTPTPTPTPTPTPTPT
Reptiles, Amphibians, Fish and Aquatic Special Concern 272829303132333435363738394041Scientific NameLeptodea ochraceaNotropis heterolepisAmia calvaCrotalus horridusCulaea inconstansEmydoidea blandingiiIchthyomyzon bdelliumLampetra aepypteraNocomis biguttatusPlestiodon laticepsUmbra limiUmbra pygmaeaAlasmidonta marginataAlasmidonta undulataAlasmidonta varicosaAmblema plicataAnodonta implicataAnodonta suborbiculataAnodontoides ferussacianusCaecidotea franziCaecidotea kenkiCaecidotea priceiCottus sp. 7Crangonyx dearolfiElliptio productaFusconaia flavaFusconaia subrotundaIchthyomyzon unicuspisLampsilis cariosaLampsilis fasciolaLampsilis radiataLasmigona complanataLasmigona compressaLasmigona subviridisLeptodea fragilisLigumia nasutaObliquaria reflexaPleurobema cordatumPleurobema sintoxiaPotamilus alatusQuadrula quadrulaCommon NameTidewater MucketBlacknose ShinerBowfinTimber RattlesnakeBrook SticklebackBlanding's TurtleOhio LampreyLeast Brook LampreyHornyhead ChubBroadhead SkinkCentral MudminnowEastern MudminnowElktoeTriangle FloaterBrook FloaterThree-ridgeAlewife FloaterFlat FloaterCylindrical PapershellFranz's Cave IsopodAn IsopodPrice's Cave IsopodCheckered SculpinPennsylvania Cave AmphipodAtlantic SpikeWabash PigtoeLong-solidSilver LampreyYellow LampmusselWavy-rayed LampmusselEastern LampmusselWhite HeelsplitterCreek HeelsplitterGreen FloaterFragile PapershellEastern PondmusselThreehorn WartybackOhio PigtoeRound PigtoePink HeelsplitterMapleleafPage 6 of 35Global ate S2S2S1S2Status CodePCPCPCPCPCPCPCPCPCPC
42434445464748495051525354Sphalloplana priceiStygobromus allegheniensisStygobromus biggersiStygobromus franziStygobromus gracilipesStygobromus pizziniiStygobromus stellmackiStygobromus tenuis potomacusToxolasma parvusTruncilla donaciformisTruncilla truncataUtterbackia imbecillisVillosa irisRefton Cave PlanarianAllegheny Cave AmphipodBiggers' Cave AmphipodFranz's Cave AmphipodShenandoah Valley Cave AmphipodPizzini's Cave AmphipodStellmack's Cave AmphipodPotomac Groundwater AmphipodLilliputFawnsfootDeertoePaper PondshellRainbow MusselPage 7 of 1SNRS1S1S1S1S1S2S1S1S3S4S1
PA Natural Heritage Program Species ListsBirds and Mammals--PA Game CommissionThreatened and Endangered Species & Special Concern SpeciesList as of Jan 28, 2014Threatened and Endangered Birds and Scientific NameArdea albaAsio flammeusBartramia longicaudaBotaurus lentiginosusChlidonias nigerCistothorus platensisCryptotis parvaEmpidonax flaviventrisFalco peregrinusGlaucomys sabrinusIxobrychus exilisLanius ludovicianus migransMyotis sodalisNyctanassa violaceaNycticorax nycticoraxRallus elegansSetophaga striataSpiza americanaSterna hirundoSciurus niger cinerusAsio otusCircus cyaneusMyotis leibiiNeotoma magisterPandion haliaetusSorex palustris punctulatusCommon NameGreat EgretShort-eared OwlUpland SandpiperAmerican BitternBlack TernSedge WrenLeast ShrewYellow-bellied FlycatcherPeregrine FalconNorthern Flying SquirrelLeast BitternMigrant Loggerhead ShrikeIndiana or Social MyotisYellow-crowned Night-heronBlack-crowned Night-heronKing RailBlackpoll WarblerDickcisselCommon TernDelmarva Fox SquirrelLong-eared OwlNorthern HarrierEastern Small-footed MyotisAllegheny WoodratOspreySouthern Water ShrewPage 8 of 35Global 5G3G3G4G5G5T3State S3BS1Status PTPTPT
Special Concern Bird and Mammal Species123456789101112131415Scientific NameCatharus ustulatusFulica americanaGallinago delicataLasionycteris noctivagansMyotis septentrionalisPodilymbus podicepsProtonotaria citreaSorex palustris albibarbisAccipiter gentilisCistothorus palustrisGallinula galeataMicrotus chrotorrhinusTyto albaArdea herodiasCommon NameSwainson's ThrushAmerican CootWilson's SnipeSilver-haired BatNorthern MyotisPied-billed GrebeProthonotary WarblerWater ShrewNorthern GoshawkMarsh WrenCommon GallinuleRock VoleBarn OwlWinter Bat ColonyGreat Blue HeronPage 9 of 35Global RankG5G5G5G5G4G5G5G5T5G5G5G5G5GNRGNRG5State B,S3NS2S3BS3BS2S2S3B,S2S3SUS3S4B,S4NStatus CodeCRCRCRCRCRCRCRCRCACACACACA
PA Natural Heritage Program Species ListsPlants, Natural Communities and Terrestrial Invertebrates--DCNRThreatened and Endangered Species & Species of ConcernList as of Jan 28, 2014Plant Threatened and Endangered 2728293031323334Scientific NameAconitum reclinatumAcorus americanusAgalinis pauperculaAlisma trivialeAlnus viridisAlopecurus carolinianusAmelanchier bartramianaAmmannia coccineaAnemone cylindricaArabis missouriensisArethusa bulbosaArnica acaulisArtemisia campestris ssp. caudataAsplenium resiliensAster borealis (name changed to Symphyotrichum boreale)Aster nemoralis (name changed to Oclemena nemoralis)Aster solidaginesus (name changed to Sericocarpus linifolius)Aster spectabilis (name changed to Eurybia spectabilis)Boltonia asteroidesCardamine pratensis var. palustrisCarex atherodesCarex aureaCarex bebbiiCarex bicknelliiCarex bullataCarex careyanaCarex collinsiiCarex crinita var. brevicrinisCarex eburneaCarex formosaCarex garberiCarex geyeriCarex gravidaCarex mitchellianaCommon NameWhite MonkshoodSweet FlagSmall-Flowered False FoxgloveBroad-Leaved Water-PlaintainMountain AlderTufted FoxtailOblong-Fruited ServiceberryScarlet AmmanniaLong-Fruited AnemoneMissouri Rock-CressSwamp-PinkLeopard’s-BaneBeach WormwoodBlack-Stemmed SpleenwortRush AsterBog AsterNarrow-Leaved White-Topped AsterLow Showy AsterAster-Like BoltoniaCuckooflowerAwned SedgeGolden-Fruited SedgeBebb’s SedgeBicknell’s SedgeBull SedgeCarey’s SedgeCollin’s SedgeShort Hair SedgeEbony SedgeHandsome SedgeElk SedgeGeyer’s SedgeHeavy SedgeMitchell’s SedgePage 10 of 35Global G5G5G5G5G4G5G4G5T5G5G4G5G50G4State 2S1S1S1S2S1S1S1S1S10S1Status PEPEPEPEPEPEPEPEPEPEPE
6061626364656667686970717273747576Carex paucifloraCarex polymorphaCarex pseudocyperusCarex retrorsaCarex typhinaCarex viridulaCerastium arvense var. villosissimum (Name changed to Cerastium velutinumvar. villossissimum)Chasmanthium laxumChenopodium foggiiChrysogonum virginianumCirsium horridulumCladium mariscoidesClematis viornaClethra acuminataClitoria marianaConioselinum chinenseCryptogramma stelleriCymophyllus fraserianusCynanchum laeveCyperus acuminatusCyperus diandrusCyperus houghtoniiCyperus refractusCyperus retrorsusCypripedium parviflorum ( Name changed to Cypripedium parviflorum var.makasin)Delphinium exaltatumDesmodium humifusumDiarrhena obovataDicentra eximiaDodecatheon meadiaDryopteris campylopteraEchinochloa walteriEleocharis caribaeaEleocharis compressaEleocharis ellipticaEleocharis obtusa var peaseiEleocharis parvulaEleocharis pauciflora var. fernaldiEleocharis quadrangulataEleocharis rostellataEleocharis tenuis var. verrucosaElephantopus carolinianusFew-Flowered SedgeVariable SedgeCyperus-Like SedgeBackward SedgeCat-Tail SedgeGreen SedgeMouse-Ear Chickweed (name changed toOctoraro Creek Chickweed)Slender Sea-OatsFogg’s GoosefootGreen-and-GoldHorrible ThistleTwig RushVase-Vine Leather-FlowerMountain Pepper-BushButterfly-PeaHemlock-ParsleySlender Rock-BrakeFraser’s SedgeSmooth Swallow-WortShort-Pointed FlatsedgeUmbrella FlatsedgeHoughton’s FlatsedgeReflexed FlatsedgeRetrorse FlatsedgeNorthern Small Yellow Lady’s-SlipperTall LarkspurTrailing Tick-TrefoilAmerican BeakgrainWild Bleeding-HeartsCommon Shooting-StarMountain Wood FernWalter’s Barnyard-GrassCapitate Spike-RushFlat-Stemmed Spike-RushSlender Spike-RushWright’s Spike-RushLittle-Spike Spike-RushFew-Flowered Spike-RushFour-Angled Spike-RushBeaked Spike-RushSlender Spike-RushElephant’s-FootPage 11 of PEPEPEPEPEPEPEPEPEPEPEPEPEPE
118Epilobium strictumEquisetum variegatumEriophorum gracileEriophorum tenellumEuphorbia ipecacuanhaeEuphorbia purpureaFestuca paradoxaFrasera caroliniensis (name changed to Swertia carolinensis )Galium labradoricumGaylussacia dumosaGeranium bicknelliiGlyceria borealisGlyceria obtusaGymnopogon ambiguusHelianthemum bicknelliiHemicarpha micrantha (name changed to Lipocarpha micrantha )Heteranthera multifloraHieracium trailliiHierochloe odorataHydrophyllum macrophyllumIodanthus pinnatifidusIris cristataIris prismaticaIris vernaIsotria medeoloidesJuncus brachycarpusJuncus dichotomusJuncus longiiJuncus militarisJuncus scirpoidesLespedeza angustifoliaLigusticum canadenseLinum intercursumLinum sulcatumListera australisListera cordataListera smalliiLithospermum carolinienseLithospermum latifoliumLobelia kalmiiLobelia puberulaLonicera oblongifoliaDowny Willow-HerbVariegated HorsetailSlender CottongrassRough CottongrassWild IpecacGlade SpurgeCluster FescueAmerican ColumboLabrador Marsh BedstrawDwarf HuckleberryCranesbillSmall-Floating Manna-GrassBlunt Manna-GrassBroad-Leaved BeardgrassBicknell’s Hoary RockroseCommon HemicarpaMulti-Flowered Mud-PlantainMaryland HawkweedVanilla Sweet-GrassLarge-Leaved Water-LeafPurple RocketCrested Dwarf IrisSlender Blue IrisDwarf IrisSmall-Whorled PogoniaShort-Fruited RushForked RushLong’s RushBayonet RushScirpus-Like RushNarrowleaf BushcloverNondo LovageSandplain Wild FlaxGrooved-Yellow FlaxSouthern TwaybladeHeart-Leaved TwaybladeKidney-Leaved TwaybladeHispid GromwellAmerican GromwellBrook LobeliaDowny LobeliaSwamp Fly HoneysucklePage 12 of PEPEPEPEPEPEPE
119 Lonicera villosa120 Ludwigia decurrens121 Ludwigia polycarpaMountain Fly HoneysuckleUpright Primrose-WillowFalse Loosestrife 36137138139140141142143Lycopodium alopecuriodes (name changed to Lycopodiella alopecuroides )Lycopodium porophilum (name changed to Huperzia porophila)Lycopus rubellusLyonia marianaMarshallia grandifloraMatelea obliquaMegalodonta beckiiMitella nudaMonarda punctataMontia chamissoiMuhlenbergia unifloraMyriophyllum exalbescens (name changed to Myriophyllum sibiricum)Myriophyllum farwelliiMyriophyllum heterophyllumMyriophyllum verticillatumNajas marinaNelumbo luteaOnosmodium molle var. hispidissimumOphioglossum engelmanniiOryzopsis pungens (name changd to Piptatherum pungens )Panicum amarum var. amarulumPanicum scoparium (name changed to Dichanthelium scoparium)Foxtail ClubmossRock ClubmossTaper-Leaved Bugle-WeedStagger-BushLarge-Flowered MarshalliaOblique MilkvineBeck’s Water-MarigoldNaked Bishop’s-CapSpotted Bee-BalmChamisso’s Miner’s-LettuceFall Dropseed MuhlyNorthern Water-MilfoilFarwell’s Water-MilfoilBroad-Leaved Water-MilfoilWhorled Water-MilfoilHolly-Leaved NaiadAmerican LotusFalse GromwellLimestone Adder’s TongueSlender Mountain-RicegrassSouthern Sea-Beach Panic-GrassVelvety 157Panicum xanthophysum (name changed to Dichanthelium xanthophysum)Parnassia glaucaPassiflora luteaPaxistima canbyiPhlox ovataPhlox subulata ssp. brittoniiPlatanthera dilatataPlatanthera hyperboreaPoa autumnalisPolemonium vanbruntiaePolygala cruciataPolygala curtissiiPolygala incarnataPolygonum careyiSlender Panic-GrassCarolina Grass-of-ParnassusPassion-FlowerCanby’s Mountain-LoverMountain PhloxMoss PinkLeafy White OrchidLeafy Northern Green OrchidAutumn BluegrassJacob’s-LadderCross-Leaved MilkwortCurtis’ MilkwortPink MilkwortCarey’s Smartweed158 Polygonum setaceum interjectum (name changed to Persicaria setacea)159 Polystichum brauniiSwamp SmartweedBraun’s Holly FernPage 13 of 5S2S1PEPE
93194195196197198199200201202203Populus balsamiferaPotamogeton friesiiPotamogeton gramineusPotamogeton hilliiPotamogeton obtusifoliusPotamogeton pulcherPotamogeton strictifoliusPotamogeton tennesseensisPotamogeton vaseyiPotentilla fruticosaPotentilla paradoxaPotentilla tridentataPrenanthes crepidineaPrunus maritimaPrunus nigraPtilimnium capillaceumPycnanthemum pycnanthemoidesPycnanthemum torreiQuercus falcataQuercus phellosQuercus shumardiiRanunculus fascicularisRhamnus lanceolataRhexia marianaRhododendron atlanticumRhynchospora capillaceaRibes missourienseRuellia humilisSagittaria calycina var. spongiosaScheuchzeria palustrisScirpus ancistrochaetusScirpus torreyi (name changed to Schoenoplectus torreyi )Scirpus acutus (name changed to Schoenoplectus acutus)Scirpus smithii (name changed to Schoenoplectus smithii)Scleria minorScleria reticularis (name changed to Scleria muhlenbergia)Scleria verticillataSedum roseaSenecio antennariifolia (name changed to Packera antennariifolia)Shepherdia canadensisSida hermaphroditaSisyrinchium atlanticumSolidago arguta var. harrisiiSolidago curtisiiBalsam P
Total Native Plants in PA Native Plants 3,348 Rare and Significant Ecological Features 415 tracked features by PNHP T&E Species Threatened plants 78 Endangered plants 228 TOTAL 306 (9% of all native plants) Special Concern Species Rare plants 39
New York Natural Heritage Program North Carolina Natural Heritage Program Ohio Natural Heritage Data Base Oklahoma Natural Heritage Inventory Ontario Natural Heritage Information Centre Oregon Natural Heritage Program Pennsylv
Heritage Local Planning Policy Framework, particularly Clause 22.05 – Heritage Policy Clause 43.01 – Heritage Overlay and Schedule to the Heritage Overlay Reference Documents – Heritage Studies 4. Methodology The scope and format of the Bayside Heritage Action Plan 2017 was informed by Heritage
STATE OF HERITAGE REVIEW Local Heritage 2020 STATE OF HERITAGE REVIEW Local Heritage 2020 Accessibility If you would like to receive this publication in an alternative format, please telephone the Heritage Council of Victoria on 9651 5060, or email heritage.council@delwp.vic.gov.au. This document is also available on the internet at
1. The World Heritage Convention and its Operational Guidelines consistently refer to World Heritage Sites as ‘Properties’ (i.e., the area of land inscribed on the World Heritage List is a “property”). The term World Heritage property is therefore used throughout this report in preference to the term World Heritage site.
World Heritage Convention and the Operational Guidelines in full. Introduction and Reader’s Guide This Resource Manual is one of a planned series of World Heritage Resource Manuals to be prepared by the UNESCO World Heritage Centre and the Advisory Bodies to the World Heritage Committee: IUCN, ICOMOS and ICCROM.
Summary 1st species: note against note 2nd species: 2 notes against 1 3rd species: 4 (or 3 notes, in ¾ time) against 1 4th species: note against note, but syncopated (suspensions) 5th species: florid counterpoint (varied rhythms, see below) Note that repeated notes are not used in species counterpoint, with one minor exception in 5th species. In the 5th species, the rhythms of the 2nd, 3rd .
Chapter 13:The origin of species Species are independently evolving lineages General lineage species concept: species are metapopulations that exchange alleles frequently enough to comprise the same gene pool Ways to identify species Biological species concept: species are
IUCN and World Heritage at a Glance Drafted World Heritage Convention with UNESCO Formal advisory body body with independent technical role to help UNESCO World Heritage Committee make informed decisions on natural heritage Evaluation of all natural and 'mixed' sites nominated for World Heritage Status, contributions to evaluations
