PSYCHE1.1011994NO.1-2Floyd Gerald Werner, 1 99 1FLOYD GERALD WERNER 192 1-1992 loyd Gerald Werner was born June 1, 192 1, to Frank and Edithmper Werner. He attended the public schools in Ottawa, Illinoishere his career in insects began with his first publication inpartment of Entomology, University of Arizona, Tucson, A Z 85721uscrim received 7 December 1993,1
2Psyche[vo .1011938, an account of the habitat and behavior of Doru aculeatum(Scudder), an earwig found in the marshes of northern Illinois. Hislove of science was fostered by a high school teacher, Charles J.Alikonis.Upon graduation Floyd enrolled at Harvard College. In 1943, hewas awarded a bachelor's degree in Biology, magna cum laude. Atthis time he was initiated into Phi Beta Kappa and Sigma Xi. During these undergraduate years Floyd served as a student assistant inthe Coleoptera Section of the Museum of Comparative Zoology,where he discovered many treasures that he would work on later inhis career.World War I1 interrupted Floyd's college career, giving him theopportunity to serve as an entomologist with the U.S. Army in theSouth Pacific. His medical survey unit worked with mosquitoes andmalaria in Okinawa and Korea. In 195 1 he returned to Okinawa atthe request of the Pacific Science Board to study sweet potato pests,as sweet potatoes were the chief source of carbohydrates for a population still in post-war recovery.In 1946-47, Floyd returned to the South Pacific as part of a scientific expedition charged with surveying the flora and fauna of thePhilippine Islands, particularly Luzon, Mindanao and Palawa. Theexpedition was sponsored by the Field Museum of Natural History,Chicago and the Philippine Bureau of Science. Collections includedeverything from bizarre mayflies in the family Prosopistomatidae togiant fruit eating bats infested with Nycteribiidae to hoodlesscobras. Stories about these times always brought a sparkle to Floyd'seyes.Returning to Harvard in 1947 after these adventures, Floydworked on a doctorate in Zoology. His specialty was insect taxonomy, in particular the Meloidae and Anthicidae. Many papers onthese families followed, climaxed by his work on the meloids ofArizona, published in 1966, and several major generic revisions ofanthicids. Floyd and fellow student William L. Nutting took longcollecting trips in the summers of 1948 and 1949 across the UnitedStates, including side trips into Canada and Mexico. They spentseveral months in the Southwest. A white bread truck, Floyd'sinfamous collecting vehicle, took him and Bill into many grandcollecting spots, including one in South Dakota that turned out tobe accessible only by a horse cart with old car tires, a surprise thatleft these compadres looking for a tow out. In 1950, Floyd was
19941Olson3awarded his Ph. D. in Zoology. His dissertation was entitled "Studies of Nearctic Anthicidae (Coleoptera)."Floyd's first job was as Assistant Professor of Zoology at theUniversity of Vermont in 1950. It was here that he met his futurewife, Frances Watson, who was also teaching in the ZoologyDepartment. This union produced three children, Susan, Williamand John. All three are commissioned law officers working forwildlife agencies in Colorado or Arizona. They are carrying ontheir father's interest in and love of the great outdoors.In 1954, Floyd returned to the scene of those monumental tripsof the late forties. He was hired by the University of Arizona entomology department, headed by Dr. Lawrence Carruth, to be part ofthe staff of the Arizona Economic Insect Survey, to expand thedepartmental insect collection as curator and to teach systematicsand other courses as needed. Collecting emphasis was on insectsimportant to the expanding agricultural community. He was promoted to Associate Professor in 1958 and Professor in 1962.At the University of Arizona, Floyd built the research collectionfrom one Cambridge Unit that held 28 Schmitt boxes into the finestSouthwest insect collection in the United States, with well overhalf a million specimens and more than thirteen thousand speciesof Arizona insects. The spectacular array of Coleoptera is the highpoint. Floyd, though, was the collector's collector, and his foraysthrough Arizona with George Butler and Bill Nutting in the earlyfifties and sixties added many representatives of Hymenoptera andDiptera, including parasites valuable for biological control. Hisresponsibilities in the department led to publications on scarabbeetles and white grubs, parasites of cotton insects and otherinsects important to agriculture in Arizona. Studies made by hisgraduate students included bee-meloid interactions, range plantinsect associations, spiders as predators in crops and various taxonomic treatments.His true love still lay with the small beetles in the familiesAnthicidae and Aderidae. In 1963 a sabbatical in South Americawith stops in Brazil and Argentina helped provide material for hiscomprehensive study of the genus Acanthinus in the New World.Other studies on anthicids included a revision of the genus Anthicus in North America and a preliminary study of the anthicids ofNew Zealand.
4Psyche[vo .101Floyd's final works included a revision of the Aderidae, a studymade possible by the excellent collections of Karl Stephan, and aspecial paper on the genus Elonus dedicated to Dr. Frank M. Carpenter, Floyd's mentor while at Harvard. Floyd was also workingon the aderids of New Zealand.After his retirement in 1989, Floyd took on the unenviable taskof Editor of the Coleopterist Bulletin, which he fulfilled to the end,his final volume of the Bulletin appearing the day after his death.He prided himself in making sure a job he took on was completedin a timely and professional manner.Because being editor of a journal and continuing work on beetleswasn't enough, Floyd accepted the challenge of writing a book onthe arthropods of the Southwest, for lay people to use to understandwhat Floyd enjoyed most, the world of insects. This book should beon the market in the spring of 1994.Floyd was a member of many professional societies and servedon a variety of their committees over the years. He was chairman ofthe Committee on Common Names of Insects and edited the 1982revision for the Entomological Society of America. He was particularly interested in the survival of taxonomy and the continuationof historical collections. He had visited a number of the collectionsof the world including those in London, Budapest and Paris, andcontributed his curatorial skills and knowledge of beetles whereverpossible at those institutions.Some of the statistics on Floyd's professional accomplishmentsinclude 100 professional papers on a variety of insects, from beetles to mantispids, parasitic wasps to spiders. His knowledge of thearthropod world was truly awesome. He described 164 species ofmeloids, anthicids, aderids and cebrionids, and three new genera inthe former three families. His name adorns such insects as an Arizona stonefly, numerous meloids, anthicids and many other taxa.As a citizen of Tucson, Floyd was active in groups such as theTucson Men's Garden Club, Tucson Cactus and Succulent Society,the Tucson Natural History Society and many others. He was askedto speak before many local groups about insects. He was an original faculty member for the Tucson Audubon Society's Desert Ecology Institute.Floyd loved to hunt and fish. He spent much of his leisure timeoutdoors experiencing the Arizona desert as a naturalist. He fishedfor sunfish and bass, bow-hunted for elk, antelope and javelina,and stalked the elusive black-tailed jackrabbit.
19941Olson5Floyd's lasting obsession was his collection of Euphorbiaceaethat occupied several greenhouses and a great part of the acre ofyard around the Werner casa. Floyd started many rare plants fromseed, priding himself in this conservation effort. He finally got tosee euphorbias first hand in the wild when he traveled to SouthAfrica with his wife and the Nuttings to attend the annual meetingof the Succulent Society in Johannesburg in 1991.It seems quite amazing that Floyd waited until retirement tosuddenly be afflicted with very untimely ills. He was huntingsquirrels the weekend before he needed his gall bladder removed.This little incident caused him to miss a couple weeks of his lastsystematics class, a rare occurrence indeed. A year later he underwent triple-bypass heart surgery, another 'minor' setback for him.He came out of those like the stout-hearted German he was, andwent back after life's challenges. His last battle, though, with cancer was a losing one and Floyd died on December 20, 1992.It seems only appropriate to end this history of my mentor, colleague and good friend with an archygram from Floyd's favoritephilosopher, Archy the Cockroach.'insects have their own pointof view about civilization a manthinks he amounts to a great dealbut to a flea or a mosquitoa human being is merely somethinggood to eat'the lives and times of archy and mehitabelby don marquisWerner, F. G. 1938. A report on the earwig Doru aculeatum (Scudder), from amarsh in Northern Illinois. Trans. Illinois State Acad. Sci. 31: 249.-- . 1943a. A revision of the genus Pleuropompha LeConte (Col., Meloidae).Psyche 50: 30-3.-- . 1943b. Three new species of Cebrio (Col., Cebrionidae). Psyche 50: 34-6.-- . 1944. Some new North American species of Epicauta (Col., Meloidae).Psyche 50: 65-73.-- . 1945. A revision of the genus Epicauta in America north of Mexico (Col.,Meloidae). Bull. Mus. Comp. Zool. 95: 421-517.
6Psyche[vo .101-- . 1948. A note on the type specimen of Bagous sellatus LeConte (Col., Curculionidae). Psyche 54: 262.Werner, F. G., and R. L. Edwards. 1948. Leptinus americanus LeConte taken on ashrew (Col., Leptinidae). Psyche 55: 5 1-4.Werner, F. G. 1949a. Epicauta diversicornis and its allies in the Neotropical Region(Col., Meloidae). Psyche 56: 74-80.-. 1949b. Additions to Epicauta, with new synonymy and a change of names(Col., Meloidae). Psyche 56: 93-1 11.-- . 1951. Additions to the Nearctic Meloidae (Col.). Psyche 57: 131-6.---. 1954a. Further notes on North American Epicauta, with new synonymy(Col., Meloidae). Psyche 60: 105-14.---. 1954b. A review of the subgenus Gnathospasta of the genus Epicauta(Col., Meloidae). Coleop. Bull. 8: 25-7.-, 1954c. Pyrota plagiata (Haag) a valid Mexican species (Col., Meloidae).Bull. Brooklyn Entomol. Soc. 49: 102-4.---. 1955. Studies in the genus Epicauta of the North American Continent(Col., Meloidae). I. The Caviceps-Group. Bull. Brooklyn Entomol. Soc. 50:1-12. 1956. Two new species of Emelinus from Arizona (Col., Aderidae). Psyche63: 30-6.Butler, G. D., and Werner, F. G. 1957. The syrphid flies associated with Arizonacrops. Arizona Agric. Exp. Sta. Tech. Bull. 132, 12 pp.Werner, F. G. 1957a. Lappus thican ormis, a new species from Michoacan. (Col.,Anthicidae). Coleop. Bull. 10: 87-9.---. 1957b. A new species of Epicauta from the Gulf Coast of Texas (Col.,Meloidae). Proc. Entomol. Soc. Washington. 59: 97-8.---. 1957c. Two cases of intestinal myiasis in man produced by Hermetia(Diptera, Stratiomyiidae). Psyche 63: 112.Werner, F. G., and Butler, G. D. 1958. The reduviids and nabids associated withArizona crops (Hemiptera). Arizona Agric. Exp. Sta. Tech. Bull. 1 33, 12 pp.Werner, F. G. 1958a. A revision of the Nearctic species of Tomoderus (Col., Anthicidae). Psyche 64: 51-9. 1958b. A new genus and species of Anthicidae from western United States(Col.). Psyche 64: 97-101.-- . 1958c. Epicauta dugesi a valid species (Col., Meloidae). Psyche 64: 107-8.---. 1958d. Some notes on Boheman's Anthicidae from "California" (Col.).Proc. Entomol. Soc. Washington 60: 2 13-1 6.---. 1959. Studies in the genus Epicauta of the North American Continent(Col.). 11. The Uniforma-Group. Coleop. Bull. 12: 1-19.Butler, G. D.; Todd, F. E.; McGregor, S. E.; and Werner, F. G. 1960. Melissodesbees in Arizona cotton fields (Hymenop., Apidae). Arizona Agric. Exp. Sta.Tech. Bull. 139, 11 pp.Werner, F. G. 1960a. A new character for the identification of the boll weevil andthe thurberi
As a citizen of Tucson, Floyd was active in groups such as the Tucson Men's Garden Club, Tucson Cactus and Succulent Society, the Tucson Natural History Society and many others. He was asked to speak before many local groups about insects. He was an origi- nal faculty member for the Tucson Audub
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